SVG sprite using CSS custom properties . Scriptura

Original SVG

<path d="" fill="#784421"/>

With default CSS variables

<path d="" fill="var(--tilion-color7, #784421)"/>

With modified CSS variables

#tilion {
  --tilion-color2: crimson;
  animation: anim-tilion 10s infinite;
}
@keyframes anim-tilion {
  25% {
    --tilion-color2: deeppink;
  }
  50% {
    --tilion-color2: mediumslateblue;
  }
  75% {
    --tilion-color2: seagreen;
  }
}